1.Municipal and Environmental Engineering School, Harbin Institute of Technology, Harbin 150086, China; Key Wood Breeding and Biotechnology Lab of China Education Ministry, Northeast Forestry University, Harbin 150024, China; 2.Municipal and Environmental Engineering School, Harbin Institute of Technology, Harbin 150086, China; 3.Key Wood Breeding and Biotechnology Lab of China Education Ministry, Northeast Forestry University, Harbin 150024, China;