¹. Lancaster Environment Centre, Lancaster University, Lancaster, LA1 4YQ, UK;². Unit of Ecology and systematics, Department of Biology, Faculty of Science, Pontificia University Javeriana, Carrera;³. No. 43-82 Ed. Jesús Emilio Ramírez (53), Bogotá, Colombia;⁴. National Park Service, Air Resources Division, Lakewood, CO 80235, USA;⁵. Synthesis Research Center of Chinese Ecosystem Research Network, Key Laboratory of Ecosystem Network Observation and Modeling, Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences, Beijing 100101, China;⁶. College of Resources and Environmental Sciences, National Academy of Agriculture Green Development, Key Laboratory of Plant-Soil Interactions of Ministry of Education, Beijing Key Laboratory of Farmland Soil Pollution Prevention and Remediation, China Agricultural University, Beijing 100193, China;⁷. School of Life Sciences, University of KwaZulu-Natal, Durban, 4041, South Africa