^a CAS Key Laboratory of Tropical Marine Bio-resources and Ecology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China
^b Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou 511458, China
^c Tropical Marine Biological Research Station in Hainan, Chinese Academy of Sciences & Key Laboratory of Tropical Marine Biotechnology of Hainan Province, Sanya Institute of Oceanology, South China Sea Institute of Oceanology, Sanya 572000, China
^d Aquatic Research Facility, Environmental Sustainability Research Centre, University of Derby, Derby DE22 1GB, UK
^e School of Ocean, Yantai University, Yantai 264005, China

^# These authors contributed equally to this work.