^a State Key Laboratory for Diagnosis and Treatment of Infectious Diseases, National Clinical Research Center for Infectious Diseases, Collaborative Innovation Center for Diagnosis and Treatment of Infectious Diseases, The First Affiliated Hospital, Zhejiang University School of Medicine, Hangzhou 310003, China
^b Jinan Microecological Biomedicine Shandong Laboratory, Jinan 250117, China
^c Department of Surgery, the First Affiliated Hospital, Zhejiang University School of Medicine, Hangzhou 310003, China
^d Key Lab of Combined Multi-Organ Transplantation, Ministry of Public Health, Hangzhou 310003, China
^e CAS Key Laboratory for Pathogenic Microbiology and Immunology, Institute of Microbiology, Chinese Academy of Sciences, Beijing 100101, China
^f University of Chinese Academy of Sciences, Beijing 100101, China
^g Shandong First Medical University & Shandong Academy of Medical Sciences, Jinan 250117, China
^h Research Units of Infectious Disease and Microecology, Chinese Academy of Medical Sciences, Hangzhou 310003, China

^# These authors contributed equally to this work.